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Achyranthes sandwicensis

(A.Gray) Di Vincenzo, Berends., Wondafr. & Borsch (2024)

This name is accepted

Kingdom: Viridiplantae Phylum: Magnoliophyta Class/Clade: Eudicot-CoreEudicot Order: Caryophyllales Family: Amaranthaceae Genus: Achyranthes

kulu‘ī [kului]

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Description

Key Characters:

Growth Form: Shrubs or small trees, branched from base or rarely a single trunk with a branched crown.

Stems: Stems (0.8–)1–4(–7) m long, 2–10 cm in diameter, silky canescent with straight, appressed hairs.

Roots:

Leaves: Leaves simple. Opposite. Blades usually elliptic, often broadly so, to ovate, broadly ovate, or lanceolate, rarely obovate, (1.5–)3–10(–13) cm long, (1–)2–4(–7.6) cm wide. Apex acuminate, acute, or obtuse, rarely retuse. Base cuneate to truncate. Surfaces usually densely pubescent with closely appressed or sometimes spreading hairs, especially on lower surface; upper surfaces usually less densely pubescent, sometimes both surfaces only sparsely pubescent or rarely glabrate. Margins entire. Pinnately veined. Petioles (3–)6–20 mm long. Stipules absent.

Flowers: Flowers in stout spikes 0.5–3(–5) cm long, 5–10 mm in diameter; peduncles (0.5–)2–5(–7.5) cm long. Flowers bisexual (perfect), lanceoloid to narrowly ovoid, actinomorphic; each flower subtended by a persistent bract and 2 bracteoles. Calyx of 4 sepals, equal, lanceolate to ovate, 2–3(–4) mm long, pubescent with long spreading hairs and with a large tuft of hairs at base, or rarely glabrous. Sepals subtended by an ovate, scarious bract ca. 1.5–2.5 mm long, ± with a spine ca. 2 mm long, usually densely pubescent, usually persistent on the rachis, bracteoles ovate to lanceolate, scarious, ca. 1.5–2 mm long, deciduous with the calyx and fruit. Corolla (petals) absent. Stamens 4, connate at base; anthers dithecal. Ovary superior, 2–3(4)-carpellate, the carpels connate to form a compound, 1-celled ovary; ovule placentation basal, free-central, or rarely apical; style 1, often lobed.

Fruit: Fruit an utricle; ca. 2 mm long; enclosed by the calyx. Seeds 1 per fruit; the surface usually lustrous; endosperm essentially absent.

Ploidy:

Habitat: Scattered to sometimes common in open dry forest; on exposed ridges; and lava fields on all of the main islands.

Elevation Range: 0–750 m.

Historical Distribution

Images

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Synonyms (36)

Nototrichium fulvum Schinz (1931)
Nototrichium sandwicense (A.Gray) Hillebr. (1888)
Nototrichium sandwicense var. decipiens Sherff (1950)
Nototrichium sandwicense var. dubium Sherff (1950)
Nototrichium sandwicense var. forbesii Sherff (1950)
Nototrichium sandwicense var. helleri Sherff (1950)
Nototrichium sandwicense var. kauaiense
Nototrichium sandwicense var. kavaiensis (A.Gray) Hillebr. (1888)
Nototrichium sandwicense var. kolekolense Sherff (1950)
Nototrichium sandwicense var. lanaiense Sherff (1950)
Nototrichium sandwicense var. lanceolatum Sherff (1950)
Nototrichium sandwicense var. latifolium Sherff (1950)
Nototrichium sandwicense var. leptopodum O.Deg. & Sherff (1950)
Nototrichium sandwicense var. longespicatum Hillebr. (1888)
Nototrichium sandwicense var. longispicatum
Nototrichium sandwicense var. macrophyllum Sherff (1950)
Nototrichium sandwicense var. mauiense O.Deg. & Sherff (1950)
Nototrichium sandwicense var. niihauense H.St.John (1959)
Nototrichium sandwicense var. olokeleanum Sherff (1950)
Nototrichium sandwicense var. pulchelloides O.Deg. & Sherff (1951)
Nototrichium sandwicense var. pulchellum Sherff (1950)
Nototrichium sandwicense var. subcordatum O.Deg. & Sherff (1950)
Nototrichium sandwicense var. syrignifolium Sherff
Nototrichium sandwicense var. syringifolium Sherff (1950)
Nototrichium viride Hillebr. (1888)
Nototrichium viride var. kalaupapae H.St.John (1980)
Nototrichium viride var. oblongifolium Sherff (1950)
Nototrichium viride var. oblongofolium Sherff
Nototrichium viride var. subtruncatum Sherff (1950)
Psilotrichum sandwicense (A.Gray) Seem.
Psilotrichum sandwicense var. kavaiensis (A.Gray) Drake (1886)
Psilotrichum sandwicense var. longespicatum (Hillebr.) Drake
Psilotrichum sandwicherse Seem. (1867)
Psilotrichum viride (Hillebr.) Drake (1892)
Ptilotus sandwicensis A.Gray (1867)
Ptilotus sandwicensis var. kavaiensis A.Gray (1867)

Uses and Culture

USES

PROPAGATION/CULTIVATION

  • The fruit of Nototrichium sandwicense are very small, about 1/16 inch in diameter and reddish. Nototrichium sandwicense is easily propagated from seeds, but it takes longer than propagating from cuttings. Sow the seeds on the surface of a mix of 1 part perlite to 1 part potting mix and water every other day. The NTBG publication (1992) states that seeds germinate in about three months. On the other hand, Lilleeng-Rosenberger's notes indicate that for one batch of fresh seed with no pretreatment 33% of the seed germinated within 1 week. (Lilleeng-Rosenberger 1996; NTBG 1992; Wagner 1990)
  • Nototrichium sandwicense grows well from cuttings. Use softwood cuttings and plant them immediately after collection. Cut off most of the upper leaves or cut them in half to reduce transpiration. Rooting hormone is not necessary, but will shorten rooting time.
  • Use a well-drained rooting medium such as clean sand, perlite, or vermiculite. NTBG suggests a mix of 3 parts perlite to 1 part vermiculite or potting soil. A humidity chamber or mist bed will speed up rooting, but is not necessary. Boche reports 50% success with stem cuttings grown in 50% shade using a medium of 3 parts cinder to 1 part perlite. Place the cuttings in the shade and water regularly. Rooting will take place in "several weeks" according to NTBG. (Boche 1992; Bornhorst 1996; Koob 1998; NTBG 1992) [Data from Herring, E. C., & Criley, R. A. (2003). The Hawaiian Native Plant Propagation Web Site: Developing a Webbased Information Resource. HortTechnology, 13(3), 545-548. https://www.ctahr.hawaii.edu/hawnprop/]

Natural History

Statewide Status

Endemic

Island Status

Ni'ihau(Incl. Lehua) Endemic
Kaua'i Endemic
O'ahu Endemic
Molokai Endemic
Lana'i Endemic
Kaho'olawe Not in flora
Maui Endemic
Hawai'i Endemic

Dispersal Agents


Pollinators

Specimens

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Bibliography

Name Published In: Taxon 73: [19 of 35] (2024)

Other References

Wagner et al. 1990:194 (Ni, K, O, Mo, L, M, Ka, H [as Nototrichium sandwicense]); Staples & Herbst 2005:99 (KEY), 103 (DESCR); Iamonico 2018:150 (N. viride Hillebr. chosen as new type, claim that N. sandwicense is a nom. nud., not followed); Di Vincenzo et al. 2024 (COMBNOV, Syn. N. sandwicense = A. sandwicensis); Faccenda, Arthur, et al. 2025:151 (remove Ka [no evidence of presence])

Occurrences

SNo. Scientific Name Locality Habitat Basis of Record Description Recorded By Record Number Island Source Date
1 Achyranthes sandwicensis Manawainui, S slope of Haleakala, on side of a gulch Side of gulch Preserved_Specimen Shrub, densely branched, 3 ft. high, looking like a Sida. Branches somewhat silvery gray. Leaves cordate, silvery pubescent below, light green with a suggestion of yellow above. Infl. short cylindrical; ovary superior, white. Only one seen. Forbes, C.N. 1816 Maui BISH 1920-03-03
2 Achyranthes sandwicensis near Crystal Gulch steep talus bank population receiving mainly indirect light, most plants sterile Preserved_Specimen vigorous bush Takeuchi, W.N. 3867 Oahu BISH 1988-05-21
3 Achyranthes sandwicensis Milolii Valley, near mouth Near mouth of valley. Preserved_Specimen Corn, C. Kauai BISH 1984-07-16
4 Achyranthes sandwicensis midway between Camp Erdman and Kaena Point, W of most prominent deep gulch, near top of cliff, just below prominent rock face windblown, dry terrain; with Artemisia, Bidens, Sicyos, Myoporum, Canthium, Sida, grass Preserved_Specimen ~2 ft. Obata, J.K. 293 Oahu BISH 1976-04-04
5 Achyranthes sandwicensis Olokele Canyon On rocks. Preserved_Specimen Rock, J.F.C. 5422 Kauai BISH 1909-09-01
6 Achyranthes sandwicensis East Maui, Makawao District, above Kihei, Puu o Kali cone aa lava; with Erythrina sandwicensis lowland dry forest with Reynoldsia sandwicensis, Nesoluma polynesicum, Sida fallax, Chenopodium oahuense, Achyranthes splendens, Diospyros sandwicensis, Rauvolfia sandwicensis, Chamaesyce, Myoporum, Capparis, Acacia koa, Prosopis Preserved_Specimen 5-7', with fruit Perlman, S. 6694 Maui BISH 1987-10-28
7 Achyranthes sandwicensis Kaupulehu Forest, between Hoolehua and Puuwaawaa with Diospyros ferrea sandwicensis, Santalum, Rauwolfia, Colubrina, Kokia Preserved_Specimen to 8' tree, branched from base Carlquist, S. 2079 Hawaii BISH 1966-07-29
8 Achyranthes sandwicensis Kapua dry, rough lava Preserved_Specimen 4 m scraggly tree MacDaniels, L.H. 266 Hawaii BISH 1926-11-16
9 Achyranthes sandwicensis Kaena, Waialua, floor of Manini Gulch with Leucaena, Panicum torridum, Plumbago zeylanicum Preserved_Specimen to 1.5 m shrub; leaves subtruncate, densely silver tomentose below, sparsely so above Nagata, K.M. 1628 Oahu BISH 1978-04-25
10 Achyranthes sandwicensis Waimea District. Koaie, dividing ridge and escarpment between Hipalau and Kawaiiki. Mixed mesic forest, summer dry, steep north facing slopes and cliff habitat w/ Diospyros sandwicensis, Acacia koaia, Planchonella sandwicensis, Santalum pyrularium, Chrysodracon aurea, Myrsine laniaensis, Rauvolfia sandwicensis Psydrax odorata, Higiscus waimeae, Flueggea neo Preserved_Specimen Tree, 2 m tall, on cliff, stems brown, arching down, in flower and fruit, occasional. Wood, K.R. 18480 Kauai BISH 2020-07-22
11 Achyranthes sandwicensis East Maui, Hana, Kanoi dry scrub; rocky slope of old aa lava flow; with Lantana camara, Opuntia megacantha Preserved_Specimen arborescent shrub; 2 m tall; crown 3 m across, trunk 0.75 dm diam. Woolford, H.A. 158 Maui BISH 1955-12-27
12 Achyranthes sandwicensis 40m makai of the Mamalahoa Hwy (190) across and SW of the entrance to the Kipuka Oweowe parking turn-out. Found in a small kipuka surrounded by a`a lava. Kipuka was dominated by invasive fountain grass (Pen set) and Lantana (Lan cam), plus a few native lama trees (Dio san), and 1 Aiea tree (Not bre) slightly farther down slope. Preserved_Specimen Parsons, E.W.R. 1 Hawaii BISH 2013-12-20
13 Achyranthes sandwicensis East side of Hoolulu Valley, on Haena-Kalalau Trail Growing amid grasses on shelf of lava cliff with Psilotum, Euphorbia, Artemisia Preserved_Specimen shrub 5 ft tall Carlquist, S. 1798 Kauai BISH 1965-08-01
14 Achyranthes sandwicensis Haena talus slope between caves and road. Preserved_Specimen Bryan Jr., E.H. 628 Kauai BISH 1928-08-06
15 Achyranthes sandwicensis Manawainui, S slope of Haleakala, on side of a gulch Side of gulch Preserved_Specimen Shrub, densely branched, 3 ft. high, looking like a Sida. Branches somewhat silvery gray. Leaves cordate, silvery pubescent below, light green with a suggestion of yellow above. Infl. short cylindrical; ovary superior, white. Only one seen. Forbes, C.N. 1816 Maui BISH 1920-03-03
16 Achyranthes sandwicensis Puuwaawaa arid aa Preserved_Specimen Degener, O. 27802 Hawaii BISH 1961-07-11
17 Achyranthes sandwicensis Waimea District, Koaie, dividing ridge between Hipalau and Kawaiiki, mixed mesic forest, summer dry, steep slopes and cliff habitat Diospyros sandwicensis, Acacia koaia, Planchonella sandwicensis, Santalum pyrularium, Dracaena aurea, Myrsine lanaiensis, Rauvolfia sandwicensis, Psydrax odorata, Hibiscus waimeae, Flueggea neowawraea Preserved_Specimen shrub, 2 m tall, very bush, silver green leaves, in early flower, observed with Panicum beecheyi, Euphorbia celastroides hanapepeensis, Panicum lineale, Eragrostis variabilis, Sicyos lanceoloidea Wood, K.R. 15362 Kauai BISH 2013-01-24
18 Achyranthes sandwicensis Na Pali Coast, Nakeikionaiwi valley diverse mesic cliff; Munroidendron racemosum, Eragrostis variabilis, Wilkesia gymnoxiphium, Schiedea apokremnos, Lobelia niihauensis, Artemisia, Wilkesia hobdyi, Peucedanum sandwicensis, Leucaena leucocephala, Schinus Preserved_Specimen Wood, K.R. 5309 Kauai BISH 1996-06-18
19 Achyranthes sandwicensis Halemanu Stream near brink of Waimea Canyon rich, wet fern-covered slope Preserved_Specimen Greenwell, A.B.H. 21513 Kauai BISH 1952-01-09
20 Achyranthes sandwicensis Waimea District, Koaie Valley, about 3/4 miles above Waimea River On cliff faces Preserved_Specimen Common shrub with gray-green foliage Herbst, D.R. 2471 Kauai BISH 1972-06-03
21 Achyranthes sandwicensis W Waimea Canyon, above Keanapahu crevices of steep slope Preserved_Specimen herbage gray, hairy St.John, H. 22974 Kauai BISH 1947-12-23
22 Achyranthes sandwicensis Waimea District. Makaha Valley. Diospyros-Acacia-Metrosideros montane mesic forest with Diospyros sandwicensis, D. hillebrandii, Alphitonia ponderosa, Pisonia sandwicensis, Dodonaea viscosa, Pleomele aurea, Psychotria mariniana, Kadua affinis, Coprosma waimeae, C. foliosa, Myrsine lan Preserved_Specimen Tree, 3.5 meters tall, stems gray, in flower. Wood, K.R. 15996 Kauai BISH 2014-07-08
23 Achyranthes sandwicensis N Kona, Puuwaawaa, 0.5 mi mauka of Mamalahoa Hwy. with Colubrina, Diospyros, Lantana Preserved_Specimen to 6' tree; leaves opposite, ovate to elliptic, base obtuse with petioles to 1 cm long; leaf blades tomentose top and bottom, below whitish Spence, G. 20 Hawaii BISH 1972-02-06
24 Achyranthes sandwicensis N. Kona, Puuwaawaa Frequent shrub along highway. growing with Metrosideros on `a`a substrate. Preserved_Specimen Ht. 5 ft. Davis, J. 514 Hawaii BISH 1981-06-14
25 Achyranthes sandwicensis NE of Kanaio, along main road aa lava kipuka Preserved_Specimen 6' strict bush Degener, O. 21982 Maui BISH 1952-02-27
26 Achyranthes sandwicensis Na Pali Coast, Nakeikionaiwi valley diverse mesic cliff; Munroidendron racemosum, Eragrostis variabilis, Wilkesia gymnoxiphium, Schiedea apokremnos, Lobelia niihauensis, Artemisia, Wilkesia hobdyi, Peucedanum sandwicensis, Leucaena leucocephala, Schinus Preserved_Specimen 3 m; many branches, multi trunked, stem gray brown, leaves green, white green below Wood, K.R. 5308 Kauai BISH 1996-06-18
27 Achyranthes sandwicensis S Kona, Ocean View on aa; with Sophora chrysophylla, Schinus, Leucaena, Lantana Preserved_Specimen 8' x 1.5 shrub, 10' spread Clarke, G. 690 Hawaii BISH 1982-04-29
28 Achyranthes sandwicensis Kawaihae; Keawewai gulch Acacia koaia forest with Diospyros, Santalum, Dodonaea, Chenopodium. Preserved_Specimen Many scattered plants 4-10 ft high with fruit. Perlman, S. 10283 Hawaii BISH 1989-08-04
29 Achyranthes sandwicensis Kona side of Puuwaawaa ranch turn-off, about 24.5 miles past Kamuela Alongside road. Substrate 'a'a; Pennisetum setaceum common. Preserved_Specimen Shrub 5 ft, 3 ft spread, excellent vigor. Scattered patches, large flrs on inflorescence. Clarke, G. 676 Hawaii BISH 1982-03-16
30 Achyranthes sandwicensis Wawaeku Gulch, E side In native remant with Lama; Olopua; Ohe makai. Preserved_Specimen 3.5 m. tree spindly with drooping branches. Hobdy, R.W. 2202 Lanai BISH 1984-10-23
31 Achyranthes sandwicensis near Crystal Gulch steep talus bank; population receiving mainly indirect light, most plants sterile Preserved_Specimen vigorous bush Takeuchi, W.N. 3867 Oahu BISH 1988-05-21
32 Achyranthes sandwicensis Napali Coast, Hanakapiai, W of Hoalulu Valley grassy ledges of basalt cliff Preserved_Specimen 2 m shrub; leaves above green, below whitish canescent; spikes white canescent, slightly pinkish St.John, H. 25544 Kauai BISH 1955-06-18
33 Achyranthes sandwicensis South Kona, Hawaiian Ocean View Estates Common in closed Sophora chrysophylla - Nototrichium - Schinus forest with Leucaena and Lantana. `A`a Substrate. Preserved_Specimen Shrub 8' tall, 10' spread, 1 1/2 basal diameter, excellent vigor. Clarke, G. 690 Hawaii BISH 1982-04-29
34 Achyranthes sandwicensis N Kona, Puuwaawaa dry aa lava Preserved_Specimen Webster, G.L. 1862 Hawaii BISH 1948-06-21
35 Achyranthes sandwicensis E Kamiloloa, NW of jeep trail walls of large gulch Preserved_Specimen Char, W.P. 82.041 Molokai BISH 1982-11-27
36 Achyranthes sandwicensis N Kona, Puuwaawaa, along Hwy. 19 on arid aa lava; with Diospyros, Metrosideros, Xylosma, Canthium, Santalum, Nototrichium, Pennisetum ruppelii Preserved_Specimen dense ~15' shrub, pale yellow green leaves Herbst, D.R. 1179 Hawaii BISH 1968-07-01
37 Achyranthes sandwicensis West Maui, behind Waikapu rock crusher Growing on the lower slope of a haole koa choked gulch in company with wiliwili, Reynoldsia, Plumbago, and Sicyos vines. Preserved_Specimen 6-8' dense shrub Hobdy, R.W. 999 Maui BISH 1981-02-24
38 Achyranthes sandwicensis Kahuku, Kau, 1887 lava flow rough aa lava Preserved_Specimen 7 m x 1 m tree St.John, H. 22420 Hawaii BISH 1946-12-27
39 Achyranthes sandwicensis Kalalau On cliff face above waterfall at beach Preserved_Specimen Milolii to 3 m. tall Hobdy, R.W. 1656 Kauai BISH 1982-08-08
40 Achyranthes sandwicensis Kaupulehu Forest Reserve, North Kona Very common in open dry forest on aa lava slope. Preserved_Specimen Small tree, 3 m. tall, grayish Fosberg, F.R. 41743 Hawaii BISH 1961-03-24
41 Achyranthes sandwicensis East Maui, Hana, Kanoi old aa lava flow; with Lantana camara, Opuntia megacantha Preserved_Specimen arborescent shrub; 2 m tall; crown 3 m across, trunk 0.75 dm diam. Woolford, H.A. 158 Maui BISH 1955-12-27
42 Achyranthes sandwicensis Na Pali Kona Trail, NW Kauai, above John Allerton's house and about 10 miles west of Hanalei, 41/2 miles from trail head, Near dry cliffs along which Brighamia insignis occurs Preserved_Specimen Gillett, G.W. 2101 Kauai BISH 1967-07-02
43 Achyranthes sandwicensis W end Dry forest Preserved_Specimen Forbes, C.N. 155 Lanai BISH 1913-06-01
44 Achyranthes sandwicensis Bottom of Kukui Trail, Waimea Canyon On sunny, arid cliffs Preserved_Specimen Erect shrub 4-7 ft. high Degener, I. 23977 Kauai BISH 1955-09-20
45 Achyranthes sandwicensis West Maui, behind Waikapu rock crusher south of the large unnamed gulch Small gulch. Growing along with other smaller kului plants with Reynoldsia and wiliwili. Preserved_Specimen 14' dense shrub, young leaves enormous Hobdy, R.W. 998 Maui BISH 1981-02-24
46 Achyranthes sandwicensis 18 mi S of Manuka Park, Kau on aa lava; with dry Metrosideros polymorpha, Dracaena aurea, Diospyros ferrea sandwicensis, Canthium odoratum, Psidium Preserved_Specimen to 10' tree Carlquist, S. 2065 Hawaii BISH 1966-07-26
47 Achyranthes sandwicensis SW of Haena; 4 1/2 miles in on Na Pali Coast Trail Among grasses on edge of dry, steep pali Preserved_Specimen shrub 2 1/2 ft. high Henrickson, J.S. 3986 Kauai BISH 1969-07-30
48 Achyranthes sandwicensis Kau, near Honuapo kipuka on ancient lava flow Preserved_Specimen Giffard, W.M. Hawaii BISH 1927-07-28
49 Achyranthes sandwicensis First valley west of Kaali Cliff Top of steep basalt talus. Preserved_Specimen Shrubs 1-2 m. tall. St.John, H. 22830 Niihau BISH 1947-08-16
50 Achyranthes sandwicensis Waimea District; Hipalau valley; North facing Native Xeric Cliffs above Native-Alien Xeric Forest; Associated Plants: Artemisia, Nototrichium sandwicense, Chamaesyce celastroides, Wilkesia gymnoxiphium, Lobelia niihauensis, Peucedanum, Bidens, Hibiscadelphus distans, Bobea timoniodes, Fleuggia, Erythrina sandwicensis, Diospyros, Myrsine, Euphorbia haeleeleana, Gahnia aspera var. globosa. Preserved_Specimen 2 m tall tree; in flower and fruit; growing at base of cliff with Hibiscadelphus distans. Wood, K.R. 1956 Kauai BISH 1992-06-06
51 Achyranthes sandwicensis Hoalulu Stream, Hanakapiai dry bush, base of cliff Preserved_Specimen 6' bush; flower spikes erect St.John, H. 13879 Kauai BISH 1933-12-30
52 Achyranthes sandwicensis N facing slope of Mahanaloa Valley Good patch of diverse native mesic forest. Diospyros spp and Euphorbia haeleleana dominant with Nestegis sandwicensis, Pisonia, Pleomele aurea and Psydrax odorata. Some Lantana camara and Grevillea also present. Preserved_Specimen Small tree 4 m tall. Silvery lvs. Fls with white, spiny bracts. Seed accessioned with Living Collections (080899). Trauernicht, C. 559 Kauai BISH 2008-08-11
53 Achyranthes sandwicensis Manuka mauka aa lava, wooded kipuka Preserved_Specimen St.John, H. 11338 Hawaii BISH 1931-12-26
54 Achyranthes sandwicensis SW Kapaka Valley Cliff ledges Preserved_Specimen Christensen, C. 175 Niihau BISH 1977-03-12
55 Achyranthes sandwicensis Halekula, Puuwaawaa, N Kona dry open forest Preserved_Specimen 6 m tree St.John, H. 18567 Hawaii BISH 1937-12-28
56 Achyranthes sandwicensis Halemanu, open valley, Open valley, rather dry Preserved_Specimen Small tree. Ht. 4 m. MacDaniels, L.H. 792 Kauai BISH 1927-02-18
57 Achyranthes sandwicensis Waimea District, Kaaweiki Ridge, just NE of (above) Polihale State Park; Disturbed dry thicket on steep slopes with Leucaena dominant, also Myoporum, Nototrichium, and native and introduced grasses Preserved_Specimen Shrub 1.5 m. tall, leaves grayish green. Infloresence whitish, stamens with yellow anthers. Lorence, D.H. 5265 Kauai BISH 1987-06-09
58 Achyranthes sandwicensis Bishop Museum full sun Preserved_Specimen 2 m bush Lau, J. 1050 Oahu BISH 1985-06-05
59 Achyranthes sandwicensis Mokouia Valley Basalt rocks, under Prosopis tree Preserved_Specimen 2 m x 2 cm shrub; leaves soft chartaceous, above dark green, below whitish green St.John, H. 23589 Niihau BISH 1949-03-30
60 Achyranthes sandwicensis Kaupulehu, on transect T12 Diospyros forest. Preserved_Specimen Subshrub 5' height, relatively common in area, most inds. fruiting. Takeuchi, W.N. 5803 Hawaii BISH 1989-05-20
61 Achyranthes sandwicensis N Kona, Hwy. 190, ~1.5 mi S of paved road to Puuwaawaa with Erythina tahitensis, Canthium odoratum, Argemone glauca, Dracaena aurea, Diospyros ferrea, Peperomia, etc.; roadside Preserved_Specimen 6-10' shrub Gustafson, R. 1082 Hawaii BISH 1978-07-13
62 Achyranthes sandwicensis Nualolo Aina Valley Growing on talus slopes near head of valley. Preserved_Specimen Small tree, 6-8' in height. ESP Field Crew 16 Kauai BISH 1979-07-18
63 Achyranthes sandwicensis Olokele Canyon On rocks. Preserved_Specimen Rock, J.F.C. 5427 Kauai BISH 1909-09-01
64 Achyranthes sandwicensis N Kona, Kona-Kohala road, near 1859 flow on old lava Preserved_Specimen 3 m bush Fosberg, F.R. 10169 Hawaii BISH 1933-09-01
65 Achyranthes sandwicensis near Crystal Gulch narrow secondary arroyo with sheer basalt walls Preserved_Specimen 6' bush Takeuchi, W.N. 3866 Oahu BISH 1988-05-21
66 Achyranthes sandwicensis N Kona, Kaupulehu, 2 mi E of Huehue dry weedy lava slope Preserved_Specimen 2 m shrub Webster, G.L. 1827 Hawaii BISH 1948-06-23
67 Achyranthes sandwicensis Na Pali Coast, Nualolo Kai, On talus slope with Lantana camara and Leucaena leucocephala. Preserved_Specimen Shrub orsmall tree Corn, C. 433 Kauai BISH 1979-07-20
68 Achyranthes sandwicensis Kumuwela ridge, Waimea dry open hillside, near stream Preserved_Specimen 6-10' bush, dense patches St.John, H. 13842 Kauai BISH 1933-12-28
69 Achyranthes sandwicensis S Kona, Ocean View on aa; with Sophora chrysophylla, Schinus, Leucaena, Lantana Preserved_Specimen 8' x 1.5 shrub, 10' spread Clarke, G. 690 Hawaii BISH 1982-04-29
70 Achyranthes sandwicensis Between Hanakapiai & Kalalau, Napali Exposed areas on cliffs in company w/Bryophyllum and grasses Preserved_Specimen Small shrub to 3' ht . Leaves, petioles, peduncles silvery green. Nagata, K.M. 1028 Kauai BISH 1972-06-11
71 Achyranthes sandwicensis N coast, Anapuhi beach, in from Huelo islet with Schiedea globosa, Pandanus, Senna gaudichaudii, Lysimachia mauritiana, Chenopodium oahuensis, Artemisia australis, Pipturus, Cyrtomium falcatum, Lythrum, Tetramolopium sylvae Preserved_Specimen with flowers and fruit Perlman, S. 6247 Molokai BISH 1987-08-01
72 Achyranthes sandwicensis Flow of 1859, N. Kona Dry lava flow Preserved_Specimen Bush 3.5m. tall Fosberg, V.O. 41 Hawaii BISH 1936-08-08
73 Achyranthes sandwicensis N Kona Distr. Puuaanahulu Ahu. Tr 46(42); N slope of Hualalai; NE Puunahaha & E of Hainoe & Kalamalu Dry exotic grassland with some native shrubs, scattered 'ohi'a and other native trees. Growing on pahoehoe Preserved_Specimen Shrub/tree; 15cm Bd; 5m tall. Monopodial form Warshauer, F.R. 2080 Hawaii BISH 1978-08-09
74 Achyranthes sandwicensis Mokuleia district, near Crystal vertical walled steam course Preserved_Specimen Paquin, D. Oahu BISH 1988-05-21
75 Achyranthes sandwicensis North Kona District. Kaupulehu Preserve, along Mamahaloa Highway (190) at 12 miles NE of Kailua Kona (by 28 mile marker) Remnant of dry forest with Diospyros sandwicensis dominant, also Colubrina, Canthium, Nototrichium, etc. with understory of introduced Pennisetum setaceum, over recent (1801) aa lava flow. Preserved_Specimen Shrub 2 m. tall, leave pale grayish green. Locally common. Lorence, D.H. 5499 Hawaii BISH 1987-08-06
76 Achyranthes sandwicensis Kukuiula, Mokuleia, Waianae Mts. (Forest trail #20a). dry forest, Sapindus oahuensis, Pleomele, Osmanthus, Erythrina, Reynoldsia, and other dryland forest trees in relict forest. Steep slope of small gulch. Preserved_Specimen 6' shrub, sterile Hatheway, W.H. 157 Oahu BISH 1950-08-08
77 Achyranthes sandwicensis Kona on recent lava flow Preserved_Specimen Beardsley, J.W. Hawaii BISH 1958-08-01
78 Achyranthes sandwicensis Na Pali Coast, Milolii Valley Growing on valley wall near streambed. Preserved_Specimen Small tree, about 6 ft. tall. ESP Field Crew 18 Kauai BISH 1979-07-19
79 Achyranthes sandwicensis Na Pali Coast State Park, Survey Supp. Sta. D, Pohakuao below Puu Ki On cliffs with Ohia, Lama, Pleomele, Hibiscus saintjohnianus, Chamaesyce, Guava Preserved_Specimen Many shrubs 4-6 ft. high with fruit Perlman, S. 10364 Kauai BISH 1989-06-14
80 Achyranthes sandwicensis Napali Coast, e. side of headland between Hanakapiai and Hoolulu Valleys Dry Scrub forest, on shelf between large boulders Preserved_Specimen Christensen, C. 45 Kauai BISH 1976-08-23
81 Achyranthes sandwicensis Halemanu Cliff lookout, Waimea, Waiahulu stream thicket, lower Acacia forest Preserved_Specimen 1-2 m shrub St.John, H. 24896 Kauai BISH 1952-12-28
82 Achyranthes sandwicensis Uao Crater, Kalaupapa, inside crater Inside crater Preserved_Specimen Harrison, B. Molokai BISH 1974-03-08
83 Achyranthes sandwicensis Waimea District, South slopes of Mahanaloa Valley South slopes Preserved_Specimen Shrub with gray-green foliage Herbst, D.R. 2830 Kauai BISH 1972-10-07
84 Achyranthes sandwicensis West Maui, behind Waikapu rock crusher, south of large unnamed gulch small gulch; steep rocky slope; with kului, Reynoldsia, wiliwili Preserved_Specimen 14' dense shrub, young leaves enormous Hobdy, R.W. 998 Maui BISH 1981-02-24
85 Achyranthes sandwicensis Na Pali-Kona Forest Reserve, Mahanaloa Valley Dry, steep N. facing slopel Associated species, relative abundance: Euphorbia, Metrosideros, Myoporum, Pelea, Pittosporum, Lantana, Kokio frequent. Preserved_Specimen Shrub 3 m. high. Gagné, W.C. 534 Kauai BISH 1970-08-22
86 Achyranthes sandwicensis Haulaula Valley Rocky streambed Preserved_Specimen Degener, O. 30268 Kauai BISH 1926-06-13
87 Achyranthes sandwicensis Napali Coast, Hanakapiai, W of Hoalulu Valley grassy ledges of basalt cliff Preserved_Specimen 2 m shrub; leaves above green, below whitish canescent; spikes white canescent, slightly pinkish St.John, H. 25544 Kauai BISH 1955-06-18
88 Achyranthes sandwicensis Na Pali Coast, Nualolo Kai Growing in old Hawaiian terrace with Leucaena leucocephala. Preserved_Specimen Small tree or shrub. Corn, C. 423 Kauai BISH 1979-07-20
89 Achyranthes sandwicensis Kona-Kohala Road near 1859 flow N. Kona on old lava Preserved_Specimen bush 3m. tall Fosberg, F.R. 10169 Hawaii BISH 1933-09-01
90 Achyranthes sandwicensis Kau, Hilea, between Hilea and Honuapo, Government Road Old aa flow, hot, dry. Preserved_Specimen Shrub 15 in. high, dbh 2 in.; Flowers with yellow stamens; straw-colored in ellipsoidal capsules; Fruit a utricle, straw-colored, enclosed in calyx. Mixed with N. sandwicense var. typicum. Russ, G.W. Hawaii BISH 1927-06-03
91 Achyranthes sandwicensis Na Pali Coast, Nualolo Kai Growing on old Hawaiian terraces and talus slopes above beach. Preserved_Specimen Small shrubby tree about 5 ft. tall. ESP Field Crew 26 Kauai BISH 1979-07-20
92 Achyranthes sandwicensis Kaupulehu Diospyros forest Preserved_Specimen subshrub 5 ft. height, relatively common in area, most inds. fruiting Takeuchi, W.N. 5803 Hawaii BISH 1989-05-20
93 Achyranthes sandwicensis Kona Road. bet lava flows of 1887 & 1907, S. Kona old lava Preserved_Specimen tree 4m. tall Fosberg, F.R. 10139 Hawaii BISH 1933-08-31
94 Achyranthes sandwicensis Waimea District; Cliffs above Polihale State Park; Haeleele Valley; North facing slope of Haeleele Ridge; Mixed Native Shrub, Coastal Dry Cliff; Associated Plants: Artemisia, Lipochaeta connata var. acris, Sida fallax, Dodonaea, Canthium, Bidens, Myoporum, Wilkesia hobdyi, Schiedea apokremnos, Peperomia, Nototrichium sandwicense. Preserved_Specimen 3 m tall tree; in flower and fruit; individual trees having both large and small leaf forms; Wood, K.R. 1950 Kauai BISH 1992-06-03
95 Achyranthes sandwicensis Kukuiula, Mokuleia, Waianae Mts (Forest trail #20a). dry forest, Sapindus oahuensis, Pleomele, Osmanthus, and many other native dryland trees on steep slopes of small gulch. Preserved_Specimen 7' shrub, much branched from base Hatheway, W.H. 168 Oahu BISH 1950-08-12
96 Achyranthes sandwicensis Kalalau trail, Napali Coast cliffs Preserved_Specimen shrub 1 m tall Stone, B.C. 827 Kauai BISH 1955-06-18
97 Achyranthes sandwicensis N Kona, Puuanahulu dry forest on aa; with Diospyros, ohia, Xylosma Preserved_Specimen Webster, G.L. 14004 Hawaii BISH 1968-06-30
98 Achyranthes sandwicensis Waialua district, Kaena, Manini Pali narrow ledges Preserved_Specimen to 5' shrub; foliage green above, silver below Herbst, D.R. 5840 Oahu BISH 1976-04-25
99 Achyranthes sandwicensis Waialua, Kaena, Manini Pali narrow ledges Preserved_Specimen to 5' shrub; foliage green above, silver below Herbst, D.R. 5840 Oahu BISH 1976-04-25
100 Achyranthes sandwicensis CGMA 1, Paliamano Gulch, above a small confluence Aspect N: At the top of a small waterfall, growing on a vertical wall. Dry riparian: Leucaena, Lantana, Cenchrus, Megathyrsus, Erythrina, Waltheria, Sida Preserved_Specimen 2 mature plants with lots of seed. Z. Pezzillo 1280 Lanai BISH 2025-10-16
101 Achyranthes sandwicensis Mauna Loa Preserved_Specimen Forbes, C.N. 3 Molokai BISH 1912-06-01
102 Achyranthes sandwicensis Awehi gulch, off Puhielelu ridge Diospyros sandwicensis, Nestegis sandwicensis, Pouteria sandwicensis, Styphelia, Metrosideros, Carex wahuensis, Gardenia brighamii, Schinus, Psidium cattleianum, Lantana camara, Psidium guajava, Erythrina sandwicensis, Rauvolfia sandwicensis, Nesoluma polynesicum, Dodonaea Preserved_Specimen Wood, K.R. 8837 Lanai BISH 2001-01-18
103 Achyranthes sandwicensis Land of Kapua, South Kona District In gulch with Lantana-Ipomoea congesta association. Preserved_Specimen Shrub to 5 m ht., with ascending branches. Nagata, K.M. 2611 Hawaii BISH 1983-02-13
104 Achyranthes sandwicensis Puu Ka Pele Forest Reserve. North slope of S fork of Polihale Ridge near oceanward end Very steep., dry slopes with Artemisia, Myoporum, Dodonaea and various weeds. Preserved_Specimen Brittle shrubs to ca. 1.5 m. high, growing on very steep, dry slopes with Aretmesia, Myoporum. Dodonaea and various weeds. Fay, J.J. 216 Kauai BISH 1989-06-14
105 Achyranthes sandwicensis Puuwaawaa arid aa Preserved_Specimen Degener, O. 27802 Hawaii BISH 1961-07-11
106 Achyranthes sandwicensis Waimea-Kohala Road, w.slopes Kohala mts. dry open forest Preserved_Specimen bush 2 m tall Fosberg, F.R. 10172 Hawaii BISH 1933-09-02
107 Achyranthes sandwicensis East Maui, Hana District, Auwahi dry scrub of Lantana, Reynoldsia, Erythrina Preserved_Specimen 3 m shrub; leaves green above, silvery below St.John, H. 25647 Maui BISH 1955-12-27
108 Achyranthes sandwicensis South Point windswept coast Preserved_Specimen Wagner, W.L. 4892 Hawaii BISH 1983-07-28
109 Achyranthes sandwicensis Halemanu Preserved_Specimen Rock, J.F.C. 1707 Kauai BISH 1909-02-14
110 Achyranthes sandwicensis Puuwaawaa road bank Preserved_Specimen to 10' shrub Stone, B.C. 3101 Hawaii BISH 1959-08-13
111 Achyranthes sandwicensis above W Manini Gulch, Waianae Mts Steep slopes & cliffs; plants @ bases of cliffs; with Canthium, Artemisia, Panicum torridum, Sida, Bidens Preserved_Specimen flowering Perlman, S. 5137 Oahu BISH 1987-01-22
112 Achyranthes sandwicensis Manawainui, S slope of Haleakala, on side of a gulch Preserved_Specimen Shrub, densely branched, 3 ft. high, looking like a Sida. Branches somewhat silvery gray. Leaves cordate, silvery pubescent below, light green with a suggestion of yellow above. Infl. short cylindrical; ovary superior, white. Only one seen. Forbes, C.N. 1816 Maui BISH 1920-03-03
113 Achyranthes sandwicensis Kaena, Manini Gulch Preserved_Specimen ~6' shrub; foliage green above, silvery beneath Herbst, D.R. 5811 Oahu BISH 1976-04-04
114 Achyranthes sandwicensis W end Dry forest Preserved_Specimen Forbes, C.N. 155 Lanai BISH 1913-06-01
115 Achyranthes sandwicensis Kaena, Manini Gulch Preserved_Specimen ~6' shrub; foliage green above, silvery beneath Herbst, D.R. 5811 Oahu BISH 1976-04-04
116 Achyranthes sandwicensis Papawai Point Preserved_Specimen Degener, O. 3341 Maui BISH 1927-07-12
117 Achyranthes sandwicensis S slope of Haleakala, near Lualailua Hills Preserved_Specimen 8' shrub, tall, multi-stemmed Nagata, K.M. 1936 Maui BISH 1979-06-07
118 Achyranthes sandwicensis Mahana near Maunalei road. Preserved_Specimen Munro, G.C. 108 Lanai BISH 1913-09-22
119 Achyranthes sandwicensis Kau Desert Preserved_Specimen Forbes, C.N. 436 Hawaii BISH 1911-08-01
120 Achyranthes sandwicensis East Molokai, Kamiloloa Preserved_Specimen Hobdy, R.W. 2356 Molokai BISH 1985-04-17
121 Achyranthes sandwicensis Honolulu, 1337 Ala Aolani, private property of J. Obata Preserved_Specimen Shrub to 2.5m. Annable, C.R. 3143 Oahu BISH 1996-11-20
122 Achyranthes sandwicensis Koloa, Lawai, NTBG, Native Hawaiian Plant Section, above road Preserved_Specimen 2 m shrub, stems erect, 2-2.5 cm.; leaves and inflorescence gray green Lorence, D.H. 8479 Kauai BISH 2000-04-26
123 Achyranthes sandwicensis Hawaii Preserved_Specimen Rémy, J. 207 Hawaii BISH 1851-01-01
124 Achyranthes sandwicensis Nakaohu Preserved_Specimen Degener, O. 19341 Maui BISH 1948-12-23
125 Achyranthes sandwicensis Kaali Cliffs, across from Puu Uala Preserved_Specimen 3 ft. Christensen, C. 158 Niihau BISH 1977-03-12
126 Achyranthes sandwicensis Puuwaawaa, last corral W of ranch house Preserved_Specimen Kondo, Y. 66 Hawaii BISH 1946-07-21
127 Achyranthes sandwicensis Waiohinu, W of 1887 flow Preserved_Specimen 15' tree St.John, H. 11032 Hawaii BISH 1931-03-12
128 Achyranthes sandwicensis N of Kohala toward Waimea, along main road 17 Preserved_Specimen Degener, O. 30271 Hawaii BISH 1976-07-31
129 Achyranthes sandwicensis North shore, East Molokai, near coast. Preserved_Specimen Shrub with white inflorescences. Anderson, S.J. 617 Molokai BISH 1983-08-25
130 Achyranthes sandwicensis Halemanu, Kapiwai Preserved_Specimen Rock, J.F.C. 2173 Kauai BISH 1909-02-14
131 Achyranthes sandwicensis Kaali Cliffs, across from Puu Uala Preserved_Specimen 4 ft. Christensen, C. 156 Niihau BISH 1977-03-12
132 Achyranthes sandwicensis Kaena, Manini Gulch Preserved_Specimen ~6' shrub; foliage green above, silvery beneath Herbst, D.R. 5811 Oahu BISH 1976-04-04
133 Achyranthes sandwicensis Puuwaawaa Preserved_Specimen Tree 15-20 feet; foliage, silvery-gray. Alexander, A.M. 5369 Hawaii BISH 1948-05-28
134 Achyranthes sandwicensis Kaena, Manini Gulch Preserved_Specimen ~6' shrub; foliage green above, silvery beneath Herbst, D.R. 5811 Oahu BISH 1976-04-04
135 Achyranthes sandwicensis Oahu Preserved_Specimen Rémy, J. 207 Oahu BISH 1854-01-01
136 Achyranthes sandwicensis Puuwaawaa Preserved_Specimen Forbes, C.N. 57 Hawaii BISH 1911-06-08
137 Achyranthes sandwicensis Nualola Kai fallen boulder, top of beach Preserved_Specimen St.John, H. 26672 Kauai BISH 1965-04-22
138 Achyranthes sandwicensis Kokee State Park; 4 to 4.5 miles along Haena-Kalalau Trail (just before leaving Hoalulu Valley, Kalalau side), above trail Preserved_Specimen Small, few-branched shrub with pale green leaves, white tomentose below; Herbst, D.R. 1101 Kauai BISH 1968-05-01
139 Achyranthes sandwicensis Mauna Loa Preserved_Specimen Tree 10 ft. high Forbes, C.N. 3 Molokai BISH 1912-06-01
140 Achyranthes sandwicensis Kaali Cliffs, across from Puu Uala Preserved_Specimen 3 ft. Christensen, C. 157 Niihau BISH 1977-03-12
141 Achyranthes sandwicensis Makolelau gulch, below Puu Kolekole with Dodonaea, Styphelia, Vaccinium, Psychotria, Diospyros, Sophora chrysophylla, Neraudia sericea, Lipochaeta rockii, Bidens menziesii, Myoporum, Wilkstroemia, Metrosideros, Pleomele, Pittosporum, Chamaesyce, Lysimachia, Viola, Pipturus, Scaevola, Urera glabra, Lantana, Melinis minutiflora, Schiedea Preserved_Specimen Perlman, S. 6660 Molokai BISH 1987-10-23
142 Achyranthes sandwicensis Kaena, Manini Gulch Preserved_Specimen shrub; foliage green above, silvery beneath Obata, J.K. 5811 Oahu BISH 1976-04-04
143 Achyranthes sandwicensis Settlement Trail Preserved_Specimen Forbes, C.N. 43 Molokai BISH 1912-06-01
144 Achyranthes sandwicensis Kona, Puuwaawaa Preserved_Specimen Rock, J.F.C. 12973 Hawaii BISH 1917-08-01
145 Achyranthes sandwicensis Puuwaawaa, near last corral W of ranch house Preserved_Specimen no flower or seed Kondo, Y. Hawaii BISH 1946-07-21
146 Achyranthes sandwicensis Kau Desert Preserved_Specimen Forbes, C.N. 436 Hawaii BISH 1911-08-01
147 Achyranthes sandwicensis Settlement Trail Preserved_Specimen Forbes, C.N. 43 Molokai BISH 1912-06-01
148 Achyranthes sandwicensis Puuwaawaa Preserved_Specimen Rock, J.F.C. 12973 Hawaii BISH 1917-08-01
149 Achyranthes sandwicensis Kalalau Valley Preserved_Specimen Forbes, C.N. 49 Kauai BISH 1909-07-17
150 Achyranthes sandwicensis Maunalei Gorge Preserved_Specimen Rock, J.F.C. 8117 Lanai BISH 1910-07-30
151 Achyranthes sandwicensis Napali Trail between Makana & Waiahuakua Preserved_Specimen Degener, O. 27790 Kauai BISH 1939-12-24
152 Achyranthes sandwicensis Mauna Loa Preserved_Specimen Forbes, C.N. 3 Molokai BISH 1912-06-01
153 Achyranthes sandwicensis Hanapepe Preserved_Specimen Mann, H. 590 Kauai BISH 1864-01-01
154 Achyranthes sandwicensis Hawaii Preserved_Specimen Rémy, J. 2071 Hawaii BISH 1851-01-01
155 Achyranthes sandwicensis Kaholuamanoa, above Waimea Preserved_Specimen Heller, A.A. 2831 Kauai BISH 1895-09-10
156 Achyranthes sandwicensis Kahalui, Maui Zoo and Botanical Garden Preserved_Specimen 2.5 m Nagata, K.M. 3199 Maui BISH 1985-03-13
157 Achyranthes sandwicensis Manini Gulch Preserved_Specimen McCauley, R. 2405 Oahu BISH 2025-07-22
158 Achyranthes sandwicensis Hawaii Preserved_Specimen Ozaki, E.T. 1369 Hawaii BISH 1954-12-31
159 Achyranthes sandwicensis Olokele Canyon Preserved_Specimen Rock, J.F.C. 5422 Kauai BISH 1909-09-01
160 Achyranthes sandwicensis S. Kona, road to Kau Preserved_Specimen Skottsberg, C.J.F. 610 Hawaii BISH 1922-09-21
161 Achyranthes sandwicensis Kalalau Trail Preserved_Specimen Lamoureux, C.H. 736 Kauai BISH 1955-06-18
162 Achyranthes sandwicensis N Kona, Puuwaawaa Preserved_Specimen Rock, J.F.C. 3814 Hawaii BISH 1909-06-14
163 Achyranthes sandwicensis Olokele Canyon Preserved_Specimen Degener, O. 30266 Kauai BISH 1926-07-03
164 Achyranthes sandwicensis N Kona, Puuwaawaa Preserved_Specimen Rock, J.F.C. 3814 Hawaii BISH 1909-06-14
165 Achyranthes sandwicensis Kaholuamanoa, above Waimea Preserved_Specimen Heller, A.A. 2831 Kauai BISH 1895-09-10
166 Achyranthes sandwicensis Kalalau Trail Preserved_Specimen Lamoureux, C.H. 731 Kauai BISH 1955-06-18
167 Achyranthes sandwicensis Kapua, Kona Preserved_Specimen Forbes, C.N. 371 Hawaii BISH 1911-07-26
168 Achyranthes sandwicensis East Molokai, Kamiloloa Preserved_Specimen Hobdy, R.W. 2354 Molokai BISH 1985-04-17
169 Achyranthes sandwicensis Between Puuwaawaa and Huehue Preserved_Specimen Degener, O. 30267 Hawaii BISH 1926-08-15
170 Achyranthes sandwicensis Waimea Preserved_Specimen Faurie, A. 1070 Kauai BISH 1910-03-01
171 Achyranthes sandwicensis Koaloha camp Preserved_Specimen Rock, J.F.C. 17142 Kauai BISH 1919-04-01
172 Achyranthes sandwicensis Waiahewahewa Gulch Preserved_Specimen Degener, O. 9869 Molokai BISH 1928-04-18
173 Achyranthes sandwicensis Puu Kolekole Preserved_Specimen Forbes, C.N. 179 Molokai BISH 1912-07-01
174 Achyranthes sandwicensis Mauna Loa Preserved_Specimen Forbes, C.N. 3 Molokai BISH 1912-06-01
175 Achyranthes sandwicensis Manawainui Preserved_Specimen Forbes, C.N. 1816 Maui BISH 1920-03-03
176 Achyranthes sandwicensis Kohala Preserved_Specimen Degener, O. 9873 Hawaii BISH 1926-08-13
177 Achyranthes sandwicensis Hanapepe Preserved_Specimen Mann, H. 590 Kauai BISH 1864-01-01
178 Achyranthes sandwicensis Paomai Preserved_Specimen Munro, G.C. 41 Lanai BISH 1914-11-18
179 Achyranthes sandwicensis Oahu Preserved_Specimen Ozaki, E.T. 1305 Oahu BISH 1954-12-23
180 Achyranthes sandwicensis Papawai Point Preserved_Specimen Degener, O. 3341 Maui BISH 1927-07-12
181 Achyranthes sandwicensis Papiloi, S slope of Haleakala Preserved_Specimen Forbes, C.N. 2081 Maui BISH 1920-03-23
182 Achyranthes sandwicensis N Kona Preserved_Specimen Alexander, A.M. 5369 Hawaii BISH 1948-05-28
183 Achyranthes sandwicensis Mauna Loa Preserved_Specimen Forbes, C.N. 3 Molokai BISH 1912-06-01
184 Achyranthes sandwicensis W end Preserved_Specimen Forbes, C.N. 155 Lanai BISH 1913-06-01
185 Achyranthes sandwicensis W Waimea drainage basin Preserved_Specimen Forbes, C.N. 1075 Kauai BISH 1917-07-03
186 Achyranthes sandwicensis Kona-Kau boundary, 1907 lava flow Preserved_Specimen Degener, O. 30269 Hawaii BISH 1976-07-25
187 Achyranthes sandwicensis Along the Hanapepe River, near the falls Preserved_Specimen Heller, A.A. 2426 Kauai BISH 1895-06-24
188 Achyranthes sandwicensis 200 yds NE of Kapaka Valley, near Kaali cliffs Preserved_Specimen Christensen, C. 174 Niihau BISH 1977-03-12
189 Achyranthes sandwicensis N Kona, Mamaloaha Hwy., mile 28, Kaupulehu Preserve Preserved_Specimen Flynn, T.W. 1263 Hawaii BISH 1985-10-15
190 Achyranthes sandwicensis along main road 19 miles from Waimea toward Kona Preserved_Specimen Degener, O. 30272 Hawaii BISH 1926-08-18
191 Achyranthes sandwicensis Settlement Trail Preserved_Specimen Forbes, C.N. 43 Molokai BISH 1912-06-01
192 Achyranthes sandwicensis S slope of Haleakala, Auwahi-Kanaio Preserved_Specimen Medeiros, A.C. 558 Maui BISH 1983-10-04
193 Achyranthes sandwicensis Waimea Preserved_Specimen Faurie, A. 1070 Kauai BISH 1910-03-01
194 Achyranthes sandwicensis Mahana Preserved_Specimen Rock, J.F.C. Molokai BISH 1910-02-01
195 Achyranthes sandwicensis below Puuwaawaa Ranch, near highway Preserved_Specimen Gillett, G.W. 1983 Hawaii BISH 1966-10-08
196 Achyranthes sandwicensis On headland between Hoolulu and Waiahuakua Valley Preserved_Specimen Christensen, C. 8 Kauai BISH 1976-05-08
197 Achyranthes sandwicensis Maunalei Gorge Preserved_Specimen Rock, J.F.C. 8117 Lanai BISH 1910-07-30
198 Achyranthes sandwicensis Oahu Preserved_Specimen Rémy, J. 207 Oahu BISH 1854-01-01
199 Achyranthes sandwicensis Kaholuamanoa Preserved_Specimen Heller, A.A. 2831 Kauai BISH 1895-09-24
200 Achyranthes sandwicensis 200 yds NE of Kapaka Valley, near Kaali cliffs Preserved_Specimen Christensen, C. 174 Niihau BISH 1977-03-12
201 Achyranthes sandwicensis Puuwaawaa, last corral W of ranch house Preserved_Specimen Kondo, Y. 66 Hawaii BISH 1946-07-21
202 Achyranthes sandwicensis 17 mi from Kohala toward Waimea Preserved_Specimen Degener, O. 30265 Hawaii BISH 1976-08-13
203 Achyranthes sandwicensis Olokele Canyon [in middle & inner part] Preserved_Specimen Skottsberg, C.J.F. 1055 Kauai BISH 1922-10-31
204 Achyranthes sandwicensis S slope of Haleakala, Auwahi-Kanaio Preserved_Specimen Medeiros, A.C. 558 Maui BISH 1983-10-04
205 Achyranthes sandwicensis near Punaluu Preserved_Specimen Degener, O. 30270 Hawaii BISH 1976-07-25
206 Achyranthes sandwicensis Upper Koaie River Preserved_Specimen Hobdy, R.W. 104 Kauai BISH 1969-07-25
207 Achyranthes sandwicensis Settlement Trail Preserved_Specimen Forbes, C.N. 43 Molokai BISH 1912-06-01
208 Achyranthes sandwicensis 0.2 mi N of Ocean View General Store, along Hwy. 11, Kahuku, Kau Preserved_Specimen Kimura, B. 4 Hawaii BISH 1980-06-29
209 Achyranthes sandwicensis Paomai Preserved_Specimen Munro, G.C. 218 Lanai BISH 1927-09-15
210 Achyranthes sandwicensis Olokele Valley Preserved_Specimen Lydgate, J.M. 4076 Kauai BISH
211 Achyranthes sandwicensis Kona, Puuwaawaa Preserved_Specimen Rock, J.F.C. 12973 Hawaii BISH 1917-08-01
212 Achyranthes sandwicensis Molueu, ridge above Manele Preserved_Specimen Munro, G.C. 192 Lanai BISH 1913-11-03
213 Achyranthes sandwicensis Kaholuamanu, Olokele Preserved_Specimen Rock, J.F.C. 17110 Kauai BISH 1916-10-01
214 Achyranthes sandwicensis Preserved_Specimen McCauley, R. 2406 Oahu BISH 2025-07-22
215 Achyranthes sandwicensis Puuwaawaa Preserved_Specimen Austin, C.F. Hawaii BISH 1912-01-01
216 Achyranthes sandwicensis near Kaena Pt Preserved_Specimen Ford, E.D. Oahu BISH 1955-04-25
217 Achyranthes sandwicensis Kawaihapai Preserved_Specimen Russ, G.W. Oahu BISH 1932-03-01
218 Achyranthes sandwicensis Kauai Preserved_Specimen Rock, J.F.C. Kauai BISH 1916-10-01
219 Achyranthes sandwicensis Kipuka Puaulu Preserved_Specimen Rock, J.F.C. Hawaii BISH 1940-01-01
220 Achyranthes sandwicensis Mahana, Mauna Loa Preserved_Specimen Rock, J.F.C. Molokai BISH 1910-02-01
221 Achyranthes sandwicensis Milolii Beach Preserved_Specimen Corn, C. Kauai BISH 1984-07-16
222 Achyranthes sandwicensis Mahana Preserved_Specimen Rock, J.F.C. Molokai BISH 1910-02-01
223 Achyranthes sandwicensis Uao Crater, Kalapaupa Preserved_Specimen Harrison, B. Molokai BISH 1974-03-08
224 Achyranthes sandwicensis Mahana Preserved_Specimen Rock, J.F.C. Molokai BISH 1910-02-01
225 Achyranthes sandwicensis Papohaku Preserved_Specimen Munro, G.C. Molokai BISH 1920-01-01
226 Achyranthes sandwicensis Honopu Beach Preserved_Specimen Davis, J. Kauai BISH 1984-07-19
227 Achyranthes sandwicensis Kauko, 6 mi from turnoff Preserved_Specimen Woolford, H.A. Hawaii BISH 1954-12-20
228 Achyranthes sandwicensis Kau, Hilea Preserved_Specimen Rock, J.F.C. Hawaii BISH 1911-04-01
229 Achyranthes sandwicensis Kauai Preserved_Specimen Rock, J.F.C. Kauai BISH 1916-10-01
230 Achyranthes sandwicensis Ulupalakua, E Maui, Makawao District Preserved_Specimen Wilder, G.P. Maui BISH 1913-01-01
231 Achyranthes sandwicensis Kawaihapae Preserved_Specimen Russ, G.W. Oahu BISH
232 Achyranthes sandwicensis Huehue Preserved_Specimen Meebold, A. Hawaii BISH 1932-05-01
233 Achyranthes sandwicensis Kauai Preserved_Specimen U.S. Exploring Expedition Kauai BISH 1840-01-01
234 Achyranthes sandwicensis Kawaihae iuka Preserved_Specimen (Not on sheet) Hawaii BISH
235 Achyranthes sandwicensis W Waimea drainage basin Preserved_Specimen Forbes, C.N. 872 Kauai BISH
236 Achyranthes sandwicensis Maunalei Gorge Preserved_Specimen Rock, J.F.C. 8117 Lanai BISH 1910-07-30
237 Achyranthes sandwicensis Hanapepe Preserved_Specimen Mann, H. 590 Kauai BISH 1864-01-01
238 Achyranthes sandwicensis Lanai Preserved_Specimen Forbes, C.N. 504 Lanai BISH 1917-09-01
239 Achyranthes sandwicensis Kaholuamanoa Preserved_Specimen Heller, A.A. 2831 Kauai BISH 1895-09-24
240 Achyranthes sandwicensis Mauna Loa Preserved_Specimen Forbes, C.N. 3 Molokai BISH 1912-06-01
241 Achyranthes sandwicensis Maunalei Gorge Preserved_Specimen Rock, J.F.C. 8117 Lanai BISH 1910-07-30
242 Achyranthes sandwicensis unknown Preserved_Specimen Hillebrand, W.B. BISH
243 Achyranthes sandwicensis Halemanu Preserved_Specimen Rock, J.F.C. 1520 Kauai BISH 1909-02-14
244 Achyranthes sandwicensis Waimea Preserved_Specimen Faurie, A. 1070 Kauai BISH 1910-03-01
245 Achyranthes sandwicensis Kaholuamanoa Preserved_Specimen Heller, A.A. 2831 Kauai BISH 1895-09-24
246 Achyranthes sandwicensis Hilea Preserved_Specimen Russ, G.W. Hawaii BISH 1927-06-03
247 Achyranthes sandwicensis Na Pali Cocast, Awa-'awapuhi Valley Rocky wall near stream at head of valley Preserved_Specimen Shrub Corn, C. 387 Kauai BISH 1979-07-19
248 Achyranthes sandwicensis Nualola Kai cliff crevices Preserved_Specimen St.John, H. 26685 Kauai BISH 1965-04-23
249 Achyranthes sandwicensis rocky ledge above mouth of Manini Gulch, Kaena, Waialua, Waianae Mts with Sida, Atriplex, Canthium, Leucaena, Eragrostis Preserved_Specimen to 2 m dense shrub; leaves densely silver-tomentose Nagata, K.M. 1636 Oahu BISH 1978-05-09
250 Achyranthes sandwicensis Olokele Canyon On rocks. Preserved_Specimen Rock, J.F.C. 5424 Kauai BISH 1909-09-01
251 Achyranthes sandwicensis Kaupulehu diverse Diospyros, Colubrina forest, loose aa clinker overgrown with fountain grass Preserved_Specimen 6' shrub, nearly all plants laden with fruit Takeuchi, W.N. 3402 Hawaii BISH 1987-02-11
252 Achyranthes sandwicensis Kamiloloa (Government) Ahupuaa, East Molokai, along 4WD road running along ridge below south fork of Kaunakakai Gulchj. Transect 3, Station 51 Mesic exotic tree community with some native trees and native and exotic grass-shrub understory. Preserved_Specimen Stemmermann, L. 3926 Molokai BISH 1979-07-15
253 Achyranthes sandwicensis Hawaii Co., Puuanahulu Game Mgmt. area, in parkland on aa flow Preserved_Specimen Occasional individual shrub to 10 ft. high, loose, open. Tomich, P.Q. 943 Hawaii BISH 1965-01-30
254 Achyranthes sandwicensis Na Pali Coast, Honopu Beach Growing on talus slope behind sand dunes. Preserved_Specimen Shrub or small tree Corn, C. 421 Kauai BISH 1979-07-20
255 Achyranthes sandwicensis Kaupulehu Forest Reserve Dry forest, part of a kipuka, growing adjacent to the Kaupuleho Lava Flow; a'a lava. Preserved_Specimen Raggedy tree, 3-4 dia., 6-8' high, leaves gray-green. Inflorescence amaranthaceous. Flowers with green perianth above, gray-green hairy beneath. Stamens bright yellow. Stern, W.L. 3123 Hawaii BISH 1974-03-28
256 Achyranthes sandwicensis Manini Pali near Manini Gulch; Kaena, Waialua, Waianae Mts., Oahu Rocky ledge overlooking mouth of Manini Gulch. Association of Sida, Atriplex, Canthium, Leucaena, Eragrostus Preserved_Specimen Dense shrub to 2 m. ht. Leaves densely silver-tomentose Nagata, K.M. 1636 Oahu BISH 1978-05-09
257 Achyranthes sandwicensis Half way between Kaalualu and Waiohinu Tree-covered dry aa lava kipuka Preserved_Specimen Degener, O. 9872 Hawaii BISH 1929-09-07
258 Achyranthes sandwicensis Moanalua Valley, home of J.Obata cultivated Preserved_Specimen 4 ft spreading shrub Takeuchi, W.N. 234 Oahu BISH 1984-07-08
259 Achyranthes sandwicensis N Kona Distr. Puuwaawaa Ahu. N slope of Hualalai, 1km E of Kileo and Potato Hill Dry koa-'ohi'a community with a native shrub-exotic grass understory. Growing on pahoehoe Preserved_Specimen Monopodial; 8cm Bd; 3m tall; 1.5m tr. len. Terrestrial rooting; 50% exposure. 1 tree seen. Jacobi, J.D. 1157 Hawaii BISH 1978-08-09
260 Achyranthes sandwicensis East Maui, above Kihei, Puu o Kali cone lava flows Preserved_Specimen 6-8' bush with tiny pubescent leaves Hobdy, R.W. 1008 Maui BISH 1981-03-05
261 Achyranthes sandwicensis Paomai dry forest Preserved_Specimen Munro, G.C. 34 Lanai BISH 1913-08-01
262 Achyranthes sandwicensis Nakaohu Aa lava waste. Preserved_Specimen Twiggy shrub 5 ft. tall. Degener, O. 19341 Maui BISH 1948-12-23
263 Achyranthes sandwicensis North Shore, East Molokai Near coast. Preserved_Specimen Shrub with white inflorescences. Anderson, S.J. 617 Molokai BISH 1983-08-25
264 Achyranthes sandwicensis Above Hanakapiai Valley, Napali Growing in crevices among stone boulders, cliffs Preserved_Specimen shrub approx. 5' height Takeuchi, W.N. 5 Kauai BISH 1983-08-17
265 Achyranthes sandwicensis Near road, by lava flow of 1907 Near road by lava flow. Preserved_Specimen Corn, C. Hawaii BISH 1977-06-27
266 Achyranthes sandwicensis Napali Coast, W Hanakoa Valley base of basalt cliff Preserved_Specimen 8 m x 1 dm tree, foliage gray green St.John, H. 23219 Kauai BISH 1947-12-31
267 Achyranthes sandwicensis Slopes of Puu Kolekole Slopes Preserved_Specimen Small tree 8 ft. tall Forbes, C.N. 179 Molokai BISH 1912-07-01
268 Achyranthes sandwicensis First valley west of Kaali Cliff Top of steep basalt talus. Preserved_Specimen Shrubs 1-2 m. tall. St.John, H. 22830 Niihau BISH 1947-08-16
269 Achyranthes sandwicensis Slopes of Puu Kolekole Slopes Preserved_Specimen Small tree 8 ft. tall Forbes, C.N. 179 Molokai BISH 1912-07-01
270 Achyranthes sandwicensis Mixed Mesic Forest. / Metrosideros, Acacia, Lepidium serra, Rumex, Hedyotis schlehtendahliana, Hedyotis foggiana, Panicum lineale, Dubautia microcephala, Ipomea tuboides, Melicope pallida, Doodia lyonii, Tectaria gaudic Preserved_Specimen 5 ft. tall. Dark reddish brown bark. Leaves silver green. Fruit w/ brn-white hairs. Tangalin, N. 999 Kauai BISH 2006-09-17
271 Achyranthes sandwicensis East Maui, Hana District, S Haleakala, Auwahi lava flows Preserved_Specimen Rock, J.F.C. Maui BISH 1954-02-07
272 Achyranthes sandwicensis Olokele Valley dryland scrub on cliffs Preserved_Specimen plants 3-5 ft. tall Christensen, C. 244 Kauai BISH 1971-06-10
273 Achyranthes sandwicensis Anapuhi to Wainene, 10-30 m from water long coastal boulder-strewn beach with vertical cliff at back; with Artemisia australis, Chenopodium oahuense, Schiedea globosa, Kalanchoe pinnata, Tetramolopium sylvae, Lythrum maritimum, Hedyotis littoralis Preserved_Specimen ~2 m shrub, many branched Wagner, W.L. 5738 Molokai BISH 1987-08-01
274 Achyranthes sandwicensis Kau, Hilea, between Hilea and Honuapo, Government Road Old aa flow, hot, dry. Preserved_Specimen Shrub 15 in. high, dbh 2 in.; Flowers with yellow stamens; straw-colored in ellipsoidal capsules; Fruit a utricle, straw-colored, enclosed in calyx. Russ, G.W. Hawaii BISH 1927-06-03
275 Achyranthes sandwicensis Waimea District, Kaaweki Ridge, just NE of (above) Polihale State Park; Disturbed dry thicket on steep slopes with Leucaena dominant, also Myoporum, Nototrichium, and native and introduced grasses Preserved_Specimen Shrub 1.5 m. tall, leaves grayish green. Infloresence whitish, stamens with yellow anthers. Lorence, D.H. 5264 Kauai BISH 1987-06-09
276 Achyranthes sandwicensis Bishop Museum full sun Preserved_Specimen 2 m bush Lau, J. 1050 Oahu BISH 1985-06-05
277 Achyranthes sandwicensis Waimea District. Koaie Canyon, about 1.mile up river from Lonomea camp. Mixed native and non-native, dry to mesic forest. On rocky, steep side of cliff above stream. W/ Aleurites moluccana, Dodonaea viscosa, Lantana camara, Melia azedarach, Kalanchoe pinnata, Artemisia australis, Psydrax odorata, Euphorbia celastroides var. hanapepensis, Hibiscus waimeae subsp, wa Preserved_Specimen Shrub about 5 feet tall. Leaves dull, light green, pubescent, involute/. Walsh, S. 526 Kauai BISH 2017-08-10
278 Achyranthes sandwicensis east Maui, Wailuku District, Keokea, SE of Pu`u O Kali. Vegetation: Erythrina, Diospyros, Sophora, Myoporum, Euphorbia, Sideroxylon, Polyscias, Nothocestrum, Sida, Chenopodium, Dodonaea, Capparis, Cocculus, Sicyos, Lipochaeta, Pellaea, Doryopteris, Ipomoea, Argemone, Waltheria, Prosopis, Melia, Leucaena, Neonotonia, Melinis, Verbesina, Lantana. Preserved_Specimen Occasional, to 4m tall. Oppenheimer, H.L. 111317 Maui BISH 2013-11-25
279 Achyranthes sandwicensis East Maui, Auwahi-Kanaio border S slope Below road. Preserved_Specimen Medeiros, A.C. 218 Maui BISH 1982-07-13
280 Achyranthes sandwicensis Kumuwela ridge, Waimea dry, steep slope Preserved_Specimen 2-3' bush, in patches St.John, H. 13850 Kauai BISH 1933-12-28
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